Near-neighbor spatial correlation (at lag 1) captures the increasingly correlation between neighboring spatial sites. It can be quantified by the spatial correlation function, or Moran’s I, between ecological states z[i, j] and z[m, n] separated by a distance r:
(1)
where is the spatial mean of the state variable (
),
is
if spatial units
and
are separated by a distance
, and is 0 otherwise,
is the total number of units separated by the distance
. In the notation
, the subscript
stands for 2-dimensional space. The near-neighbor spatial correlation
, the analog of autocorrelation at lag 1 for time series, is calculated for the distance
corresponding nearest neighboring units of the system.